I dug out the Corky Meyer's article on the Joint Fighter conference (Flight Journal WWII Fighter issue):
"The Seafire had such delightful upright flying qualities that knowing it had an inverted fuel and oil system, I decided to try inverted figure-8s. They were easy as pie, even hanging by the complicated, but comfortable, British pilotrestraint harness. I was surprised to hear myself laughing as if I were crazy. I have never enjoyed a flight in a fighter as much before or since or felt so comfortable in a plane at any flight attitude."
Anyway, the Seafire in the Conference might had had a some sort of special engine. There were two kind of lubrication systems in the Merlin and the late models got improved, so called end-to-end, oil system. That was supposed to be introduced in the Merlin 100 series (and V-1650-9 got it as well) but I don't know if it was used in the Merlin 55 of the Seafire III.