Spit VIII: 120gal, 4x .303, 2x 20mm.
Takeoff weight 7,780lb, moment +41,553 lb in
=> CG is 41,533/7,780 = 5.3" aft of D
Spit XIV: 110gal, 4x .303, 2x 20mm.
Takeoff weight 8,475lb, moment +30,699 lb in
=> CG is 30,699/8,475 = 3.6" aft of D
so CG is 1.7" further forward in the XIV. I cant imagine that this is enough to explain the vastly different handling characteristics, which is why I'm sticking with the idea of the location of the extra mass and the increased moment of inertia explaining it.