Seat position and G forces....
The relevant point here is the relationship of the heart to the head. If the head can be placed 'level' with the heart, then the heart has an easier job of pumping blood to the head...the more upright the head is, the harder it is for the heart to pump blood 'uphill', ie under G. Obviously, it's the blood in the head that is important here, not the blood that is in the legs!
There were several designs in WW2 that had the pilot in the prone position (laying on his stomach) to provide relief from G forces...none of them were operationally workable...although they did in fact reduce or slow G-induced 'blackout'.
It is doubtful that leg position had a significant impact on the head/heart issue...emphasis on the word 'significant'. It is far more likely that the desire to keep aircraft size and vertical cross-section to a minimum was more in play in the seat/rudder position issue.
Andy