I've been reading up on this problem and it was actually a little more involved. The carburetter was employed in the Spitfire in place of fuel injection because they believed it increased the effectiveness of the supercharger and hence gave more power (the Germans were forced to employ fuel injection because they went and built their engine upside down in the first place

).
The early versions used an SU carburetter. When bunting, the
fuel was forced to the top of the float chambers causing the initial fuel starvation (spluttering and loss of power). If the negative G manoeuvre was sustained then the fuel would accumulate in the top of the float chamber and push the float downwards opening the SU's needle valve to maximum, flooding the engine and shutting it down completely.
Tilly's orifice was fitted to the fuel lines which restricted the flow and allowed just enough fuel for maximum power (clever eh?) it even came in two settings, for boosted and normal engines. The needles of the carburetters were also modified and the fuel feed moved from the bottom to the side of the float chamber.
These solutions allowed the Spitty pilots to follow the dastardly Hun through his cowardly bunting escapes and fill the blighter full of 303, preferably aiming for his knockwurst
[1]. These problems were completely resolved with the introduction of the Bendix pressurised carburettors in '42.
[1] "Bag the Hun: Estimation of Range & Angle Off". Air Ministry - 1943 [http://bbs.hitechcreations.com/smf/index.php/topic,297962.0.html]